Lentic Ecosystems


Stationary or Relatively Still Water

A lake ecosystem includes biotic (living) plants, animals and micro-organisms, as well as abiotic (nonliving) physical and chemical interactions.

Lake ecosystems are a prime example of lentic ecosystems. Lentic refers to stationary or relatively still water, from the Latin lentus, which means sluggish. Lentic waters range from ponds to lakes to wetlands, and much of this article applies to lentic ecosystems in general. Lentic ecosystems can be compared with lotic ecosystems, which involve flowing terrestrial waters such as rivers and streams. Together, these two fields form the more general study area of freshwater or aquatic ecology.

Lentic systems are diverse, ranging from a small, temporary rainwater pool a few inches deep to Lake Baikal, which has a maximum depth of 1642 m. The general distinction between pools/ponds and lakes is vague, but Brown states that ponds and pools have their entire bottom surfaces exposed to light, while lakes do not. In addition, some lakes become seasonally stratified (discussed in more detail below.) Ponds and pools have two regions: the pelagic open water zone, and the benthic zone, which comprises the bottom and shore regions. Since lakes have deep bottom regions not exposed to light, these systems have an additional zone, the profundal. These three areas can have very different abiotic conditions and, hence, host species that are specifically adapted to live there.

PH – Balance


Acidification

Sulfur dioxide and nitrogen oxides are naturally released from volcanoes, organic compounds in the soil, wetlands, and marine systems, but the majority of these compounds come from the combustion of coal, oil, gasoline, and the smelting of ores containing sulfur.[3] These substances dissolve in atmospheric moisture and enter lentic systems as acid rain.[1] Lakes and ponds that contain bedrock that is rich in carbonates have a natural buffer, resulting in no alteration of pH. Systems without this bedrock, however, are very sensitive to acid inputs because they have a low neutralizing capacity, resulting in pH declines even with only small inputs of acid.[3] At a pH of 5–6 algal species diversity and biomass decrease considerably, leading to an increase in water transparency – a characteristic feature of acidified lakes. As the pH continues lower, all fauna becomes less diverse. The most significant feature is the disruption of fish reproduction. Thus, the population is eventually composed of few, old individuals that eventually die and leave the systems without fishes.[2][3] Acid rain has been especially harmful to lakes in Scandinavia, western Scotland, west Wales and the north eastern United States.


Eutrophication

Eutrophic systems contain a high concentration of phosphorus (~30 µg/L), nitrogen (~1500 µg/L), or both.[2] Phosphorus enters lentic waters from sewage treatment effluents, discharge from raw sewage, or from runoff of farmland. Nitrogen mostly comes from agricultural fertilizers from runoff or leaching and subsequent groundwater flow. This increase in nutrients required for primary producers results in a massive increase of phytoplankton growth, termed a plankton bloom. This bloom decreases water transparency, leading to the loss of submerged plants. The resultant reduction in habitat structure has negative impacts on the species’ that utilize it for spawning, maturation and general survival. Additionally, the large number of short-lived phytoplankton result in a massive amount of dead biomass settling into the sediment.[5] Bacteria need large amounts of oxygen to decompose this material, reducing the oxygen concentration of the water. This is especially pronounced in stratified lakes when the thermocline prevents oxygen rich water from the surface to mix with lower levels. Low or anoxic conditions preclude the existence of many taxa that are not physiologically tolerant of these conditions.[2]


Invasive species

Invasive species have been introduced to lentic systems through both purposeful events (e.g. stocking game and food species) as well as unintentional events (e.g. in ballast water). These organisms can affect natives via competition for prey or habitat, predation, habitat alteration, hybridization, or the introduction of harmful diseases and parasites.[4] With regard to native species, invaders may cause changes in size and age structure, distribution, density, population growth, and may even drive populations to extinction.[2] Examples of prominent invaders of lentic systems include the zebra mussel and sea lamprey in the Great Lakes.

Freshwater algal blooms


Further information: Nutrient pollution and Eutrophication

Freshwater algal blooms are the result of an excess of nutrients, particularly some phosphates.[3][4] The excess of nutrients may originate from fertilizers that are applied to land for agricultural or recreational purposes. They may also originate from household cleaning products containing phosphorus.[5] These nutrients can then enter watersheds through water runoff.[6] Excess carbon and nitrogen have also been suspected as causes. Presence of residual sodium carbonate acts as catalyst for the algae to bloom by providing dissolved carbon dioxide for enhanced photosynthesis in the presence of nutrients.

When phosphates are introduced into water systems, higher concentrations cause increased growth of algae and plants. Algae tend to grow very quickly under high nutrient availability, but each alga is short-lived, and the result is a high concentration of dead organic matter which starts to decay. The decay process consumes dissolved oxygen in the water, resulting in hypoxic conditions. Without sufficient dissolved oxygen in the water, animals and plants may die off in large numbers. Use of an Olszewski tube can help combat these problems with hypolimnetic withdrawal.

Blooms may be observed in freshwater aquariums when fish are overfed and excess nutrients are not absorbed by plants. These are generally harmful for fish, and the situation can be corrected by changing the water in the tank and then reducing the amount of food given.